Elephas hysudrindicus

Elephas hysudrindicus, also known in Indonesia as the Blora elephant, is an extinct elephant from the Pleistocene of Java. It is anatomically distinct from the Asian elephant and existed from around the end of the Early Pleistocene until the end of the Middle Pleistocene, when the modern Asian elephant replaced it on Java. It coexisted with the proboscidean Stegodon trigonocephalus and with Homo erectus. Eugene Dubois described it in 1908 without designating a holotype, and in 2017 a partial skull in the Naturalis Biodiversity Center in Leiden, excavated at Tinggang, Bojonegoro, was designated the lectotype. It is considered closely related to Elephas hysudricus of mainland Asia and possibly descended from it. A nearly complete fossil was excavated in 2009 at Sunggun near Blora. Isotopic analysis suggests a diet dominated by C4 grazing in open habitats.